Scientific Papers
Given the central role of scientific research in MRF, we have created a section for all of the peer reviewed scientific papers produced by MRF staff or papers using MRF material. Many MRF volunteers helped excavate many of the localities that are now being published on. There is a brief summary of each paper. A pdf of the paper can be downloaded by clicking on the reference.
Click on an item to read the full paper.  Turtles all the way down: loggerheads at the root of the chelonian treeEvolution and Development (2009) Tyler Lyson and Scott F. Gilbert Summary This paper discusses the amazing turtle with teeth and half a shell from China, Odontochelys semitestacea. This amazing turtle surprised the scientific community and reopened up several important questions regarding the origin of turtles, the origin of the turtle shell, and their original paleoecology (aquatic or terrestrial).  A Revision of Plesiobaena (Testudines: Baenidae) and an Assessment of Baenid Ecology Across the K/T BoundaryJournal of Paleontology (2009) Tyler R. Lyson and Walter G. Joyce Summary A new species of baenid turtle is described from the Hell Creek Formation. Material of this new species, Peckemys brinkman, was found at MRF's Turtle Graveyard locality. This locality is unusual in that it represents a true snapshot in time (i.e. it is constrained both temporally and spatially). The locality is the most diverse fossil turtle assemblage. Much of the material described in this paper was found by MRF volunteers. The paper discusses the diversity of turtles found at this locality as well as the number of baenid turtles that go extinct at the K/T boundary. Baenid turtles do surprisingly well across the boundary, with seven of nine baenids surviving into the Paleocene. Interestingly, four of the seven survivors are interpreted as having a mollusk and snail type diet.  A New Species of Palatobaena (Testudines: Baenidae) and a Maximum Parsimony and Bayesian Phylogenetic Analysis of BaenidaeJournal of Paleontology (2009) Tyler R. Lyson and Walter G. Joyce Summary This is the first of a series of papers on Marmarth Research Foundation's Turtle Graveyard and Turtle Ridge localities. These localities are the world's most diverse and largest fossil turtle quarries. They have produced a number of new species, including the one described in this paper: Palatobaena cohen. This enigmatic turtle was named after MRF board of director and longtime volunteer Steven Cohen. The paper discusses the geology of the Turtle Graveyard locality, a detailed description of the skull and shell, and a phylogenetic analysis of the group of turtle to which Pa. cohen belongs, Baenidae. Hundreds of MRF volunteers have helped excavate this locality and found the material described in this paper including John Akerly (shell), Pam Hollocher (shell), Lou Tremblay (skull and lower jaw), and Norman Gardiner (skull).  Evolution of the Respiratory System in Nonavian Theropods: Evidence from Rib and Vertebral MorphologyThe Anatomical Record (2009) Emma R. Schachner, Tyler R. Lyson, and Peter Dodson Summary This study analyzes rib and vertebral anatomy of carnivorous dinosaurs in order to help determine the type of respiratory system present in carnivorous dinosaurs. Did carnivorous dinosaurs have a flow-through ventilation system as in birds? Or did they have a more crocodilian-like hepatic-piston respiratory system? The study shows that a crocodilian-like hepatic-piston lung would be functionally and biomechanically untenable and that carnivorous dinosaurs share respiratory anatomical characters with birds (i.e. air sacs) and likely had a more bird-like respiratory system.  Mineralized soft-tissue structure and chemistry in a mummified hadrosaur from Hell Creek Formation, North DakotaProceedings of The Royal Society B (2009) Phillip L. Manning, Peter M. Morris, Adam McMahon, Emrys Jones, Andy Gize, Joe H. S. Macquaker, George Wolff, Anu Thompson, Jim Marshall, Kevin G. Taylor, Tyler Lyson, Simon Gaskell, Onrapak Reamtong, William I. Sellers, Bart E. van Dongen, Mike Buckley and Roy A. Wogelius Summary This is the first of a series of papers on the mummified dinosaur "Dakota." This paper was selected by Discover Magazine as one of the top 100 best scientific papers of 2009 (ranking 46). The paper goes through a number of geochemical analyses in search of original biomolecules from "Dakota." In addition, the paper discusses how "Dakota" was preserved and describes the 3-D preservation of the skin.  Virtual Palaeontology: Gait Reconstruction of Extinct Vertebrates Using High Performance ComputingPalaeontologia Electronica (2009) William I. Sellers, Phillip L. Manning, Tyler R. Lyson, Kent Stevens, and Lee Margetts Summary Were hadrosaur dinosaurs running on two or four legs at top speed? This paper uses computer simulations to try and address this question. An 8000 core computer was used to produce mechanically and physiologically plausible gaits and trackway patterns for an Edmontosaur annectens dinosaur. Galloping was determined to be the most plausible and fastest (16 ms-1) gait, followed by bipedal running (14ms-1).
Watch the Edmontosaur gallop (mov, 11MB), run (mov, 10.5MB), and even hop (mov, 8MB)!  Two New Plastomenine Softshell Turtles from the Paleocene of Montana and WyomingBulletin of the Peabody Museum of Natural History (2009) Walter G. Joyce, Ariel Revan, Tyler R. Lyson, and Igor G. Danilov Summary Two new species of soft-shelled (Trionychidae) turtles are described and their paleoecology discussed. The subgroup of trionychids described, Plastomenidae, are unusual in that they have a well-ossified bottom shell (plastron). The group is thought to be part of the lineage that gave rise to modern day flap shelled turtles (Cyclanorbinae).
Triceratops squamosal
Homer 2005
Marmarth Research Foundation
P.O. Box 5
Marmarth, ND 58643
Phone: (610) 937-7916
(June-Aug): (701) 279-6601
Email: tyler.lyson@mrfdigs.com
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